Checklist of Diptera of the Czech Republic and Slovakia
electronic version 2, 2009
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Scatopsidae Newman, 1834

Jean-Paul Haenni

Muséum d’histoire naturelle, rue des Terreaux 14, CH-2000 Neuchâtel, Switzerland;

The Scatopsidae are a medium-sized but still inadequately known family of derived nematocerous Diptera. Adults are minute to small midges ranging from 0.5 to 4.0 mm in length, generally dark in colour, sometimes more or less marked with yellow. Body rather strongly built, often quite elongated, with parallel-sided abdomen. Eyes holoptic, forming a characteristic supraantennal bridge. Antenna comparatively short, composed of two basal segments and a flagellum of 5-10 flagellomeres that are generally wider than long. Wing venation reduced, generally only anterior veins pigmented. Larvae flattened, with a complete cephalic capsule, peripneustic. All known larvae are saprophagous, living in a wide variety of media (rotten plants and fruits, rotten wood, under decaying bark of dead trees, soil, leaf litter, fungi, dung, etc) (Haenni & Vaillant 1994). The adults may be found in nearly all kinds of biotopes, but more frequently in open, fresh and marshy areas.

The world fauna of Scatopsidae comprises some 350 species (Haenni 1997), of which 102 species in 22 genera are known in Europe (Haenni 2007 and recent additions). 52 species are included in the present checklist: 46 from the Czech Republic (41 from Bohemia and 42 from Moravia) and 43 from Slovakia. Since the preceding PCV2 (Martinovský 1997b), which was mainly based upon the important papers by Martinovský (1997a) for the Czech Republic and by Kozánek and Roller (1997) for Slovakia, the number of species had increased by eight in the Czech Republic (ten in Bohemia and seven in Moravia) while it had remained the same in Slovakia. The rather numerous additional records may be found in a set of recent faunistic papers (Haenni & Barták 2000, 2006, Haenni et al. 2005, Martinovský 1998). On the other hand, two nomenclatorial changes have been proposed by Haenni (1998, 2002). By contrast only one new record for Slovakia has been added (Haenni 2009) since the previous electronic version (ECV1, Haenni 2006). The species richness of the scatopsid fauna is rather well known in the Czech Republic and in Slovakia, but the number of species will continue to grow as there are several new species awaiting description.

The male and female genital characters enable safe identifications to be made. A key to the Palaearctic genera (both larvae and adults) may be found in Haenni (1997), where references to papers for identification to species level are also included. The classification used here follows Amorim (1994) and Chandler (1998), while the nomenclature follows Haenni (2007).


[1] Amorim D. de Souza 1994: A new suprageneric classification of the Scatopsidae (Diptera: Psychodomorpha). Iheringia, Série Zooogia 77: 107-112.

[2] Chandler P. 1998: Scatopsidae. In Chandler P. (ed.): Checklists of Insects of the British Isles (New Series). Part 1. Diptera (Incorporating a List of Irish Diptera). Handbooks for the Identification of  British Insects, Vol. 12. Royal Entomological Society, London, pp. 47-48.

[3] Haenni J.-P. 1997: Family Scatopsidae. In Papp L. & Darvas B. (eds): Contributions to a Manual of Palaearctic Diptera (with special reference to flies of economic importance). Volume 2. Nematocera and Lower Brachycera. Science Herald, Budapest, pp. 255-272.

[4] Haenni J.-P. 1998: Scatopsidae. In: Merz B., Bächli G., Haenni J.-P. & Gonseth Y. (eds): Diptera Checklist. Fauna Helvetica 1. CSCF/ SEG, Neuchâtel, pp. 141-144.

[5] Haenni J.-P. 2002: Redescription de Scatopse filamentosa Duda, 1928, avec une nouvelle synonymie (Diptera, Scatopsidae). Bulletin de la Société Neuchâteloise des Sciences Naturelles 125: 93-96.

[6] Haenni J.-P. 2006: Scatopsidae. In: Jedlička L., Stloukalová V. & Kúdela M. (eds): Checklist of Diptera of the Czech Republic and Slovakia. Electronic version 1. <>. ISBN 80-969629-0-6. Retrieved 04.06.2009.

[7] Haenni J.-P. 2007: Scatopsidae. In Jong H. de (ed.): Fauna Europaea: Diptera, Nematocera. Fauna Europaea version 1.3. <>. Retrieved 04.06.2009.

[8] Haenni J.-P. 2009: Scatopsidae. In Roháček J. & Ševčík J. (eds): Diptera of the Poľana Protected Landscape Area – Biosphere Reserve (Central Slovakia). SNC SR, Administration of the PLA – BR Poľana, Zvolen, pp. 94-96.

[9] Haenni J.-P. & Barták M. 2000: Scatopsidae. In Barták M. & Vaňhara J. (eds): Diptera in an industrially affected region (North-Western Bohemia, Bilina and Duchcov environs), I. Folia Facultatis Scientiarum Naturalium Universitatis Masarykianae Brunensis, Biologia 104: 105-109.

[10] Haenni J.-P. & Barták M. 2006: Faunistic records from the Czech Republic (Diptera: Scatopsidae). Entomofauna carpathica 18: 85.

[11] Haenni J.-P., Barták M. & Kubík Š. 2005:  Scatopsidae. In Barták M. & Kubík Š. (eds): Diptera of Podyjí National Park and its Environs. Česká zemědělská univerzita, Praha, pp. 90-93.

[12] Haenni J.-P. & Vaillant F. 1994: Description of dendrolimnobiontic larvae of Scatopsidae (Diptera) with a review of our knowledge of the preimaginal stages of the family. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 67: 43-59.

[13] Kozánek M. & Roller L. 1997: A study of  scatopsid (Diptera: Scatopsidae) communities in Slovakia. Biologia, Bratislava 52: 637-646.

[14] Martinovský J. 1997a: Contribution to the knowledge of Scatopsidae (Diptera) from the Czech and Slovak Republics, with descriptions of two new species. Časopis Slezkého Muzea v Opave (A) 45 (1996): 193-210.

[15] Martinovský J. 1997b: Scatopsidae. In Chvála M. (ed.): Check List of Diptera (Insecta) of the Czech and Slovak Republics. Carolinum – Charles University Press, Prague, pp. 32-33.

[16] Martinovský J. 1998. Scatopsidae. In Rozkožný R. & Vaňhara J. (eds): Diptera of the Pálava Biosphere Reserve of UNESCO, I. Folia Facultatis Scientiarum Naturalium Universitatis Masarykianae Brunensis, Biologia 99: 83-85.

Aspistes Meigen, 1818
berolinensis Meigen, 1818  CZ   (B M ) SK
Ectaetia Enderlein, 1912
clavipes (Loew, 1846)  CZ   (B M ) SK
Anapausis Enderlein, 1912
albohalterata Duda, 1928  CZ   (B M ) SK
dufourella Haenni, 1984  CZ   ( M ) SK
floricola Chandler, 1999 CZ (B M )
haemorrhoidalis Duda, 1928  CZ   (B M ) SK
helvetica Haenni, 1984  CZ   (B M ) SK
inermis (Ruthé, 1831)  CZ   (B M ) SK
nigripes (Zetterstedt, 1860)  CZ   (B M )
rectinervis Duda, 1928  CZ   (B M ) SK
rohaceki Martinovský, 1997  SK
soluta (Loew, 1846)  CZ   (B M ) SK
talpae (Verrall, 1912)  CZ   (B M ) SK
COLOBOSTEMATINI in PCV2 all species in tribe SCATOPSINI [1]
Colobostema Enderlein, 1926
infumatum (Haliday, 1833) CZ (B )
nigripenne (Meigen, 1830)  CZ   (B M ) SK
obscuritarse (Strobl, 1898) CZ ( M )
triste (Zetterstedt, 1850)  CZ   (B M ) SK
Efcookella Haenni, 1998
albitarsis (Zetterstedt, 1850)  CZ   (B M ) SK in PCV2 in genus Cookella Freeman, 1985, preocc. [4]
Ferneiella Cook, 1974
brevifurca (Enderlein, 1912)  CZ   (B )
incompleta (Verrall, 1886)  CZ   (B M ) SK
Holoplagia Enderlein, 1912
bullata (Edwards, 1925)  CZ   (B M ) SK
lucifuga (Loew, 1870)  CZ   (B M ) SK
richardsi (Edwards, 1934)  CZ ( M ) SK
transversalis (Loew, 1846)  CZ   (B M ) SK
Thripomorpha Enderlein, 1905
bifida (Zilahy-Sebess, 1956)  CZ   (B M ) SK
freyi (Cook, 1969)  CZ   (B M )
haennii Martinovský, 1997  SK
halteratum (Meigen, 1838)  CZ   (B M ) SK
paludicola Enderlein, 1905  CZ   (B M ) SK
verralli (Edwards, 1934)  CZ   (B M ) SK
Apiloscatopse Cook, 1974
flavicollis (Meigen, 1818)  CZ   (B M ) SK
flavocincta (Duda, 1928)  CZ   (B M )
gracilis (Duda, 1928)  CZ   (B M ) SK
filamentosa (Duda, 1928) CZ ( M ) SK syn. hennigi Cook, 1974 [5]
mattheyi Haenni, 1981  CZ   ( M )
picea (Meigen, 1818)  CZ   (B M ) SK
scutellata (Loew, 1846)  CZ   (B M ) SK
styriaca (Enderlein, 1926)  SK
Pharsoreichertella Cook, 1974 in PCV2 as subgenus in Reichertella Enderlein, 1912 [1]
hamifera (Strobl, 1909)  CZ   (B )
simplicinervis (Duda, 1928)  SK
Reichertella Enderlein, 1912 in PCV2 all species in suppressed subgenus Reichertella Enderlein, 1912 [1]
geniculata (Zetterstedt, 1850)  CZ   (B M ) SK
nigra (Meigen, 1804)  CZ   (B M ) SK
pulicaria (Loew, 1846)  CZ   (B M ) SK
Scatopse Geoffroy, 1762
lapponica Duda, 1928 CZ (B ) SK
notata (Linné, 1758)  CZ   (B M ) SK
Coboldia Melander, 1916
fuscipes (Meigen, 1830)  CZ   (B M ) SK
Rhexoza Enderlein, 1936
subnitens (Verrall, 1886)  CZ   (B M ) SK
Swammerdamella Enderlein, 1913
acuta Cook, 1956  CZ   (B M ) SK
adercotris Cook, 1972  CZ   (B M ) SK
brevicornis (Meigen, 1830)  CZ   (B M ) SK
genypodis Cook, 1972  SK
pediculata (Duda, 1928)  SK