Checklist of Diptera of the Czech Republic and Slovakia
electronic version 2, 2009
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Scathophagidae Robineau-Desvoidy, 1830

František Šifner

V štíhlách 1311, CZ–142 00 Praha 4 – Krč, Czech Republic

Body-length 5.0-15.0 mm, ground-colour dark, black or yellow, frequently shining, entirely or partially grey dusted, strongly bristled and often yellow haired. The present status of the family is well-founded and unambiguous, with the following principal characters: lower calypter (squama alaris) greatly reduced and linear, head more or less spherical and eyes in both sexes widely separated, frontal vitta in both sexes wide and without a pair of crossed frontal bristles, second antennomere with distinct and complete dorsal longitudinal seam, venation of wing almost constant and without a strong costal spine at end of subcostal vein, katepisternum with one to three bristles, two pairs of gonapophyses. Bernasconi et al. (2000) addressed the problems of phylogeny by carrying out analyses based on COI and Cytb sequences, and they confirmed the monophyly of the Scathophagidae within the limits of the muscoid families.

The adults occur in sheltered and moist habitats of various types, forests, lowland to montane meadows, littoral vegetation, marshes, peat-bogs, etc. Adults are predaceous and hunt small insects, exceptionally consuming nectar in flowers, whilst others are coprophagous and perhaps saprophagous; several arctic species have often been observed on carrion. The eggs are attached to the surface of leaves or are inserted into plant tissues or other substrates. The larvae are phytophagous and mine in the leaves and stems particularly of monocotyledonous plants and, much more rarely, of dicotyledonous plants; other larvae are predaceous and saprophagous, living in soil or stagnant and running water; the larvae of some species occur in excrement, feeding mostly as predators. The larval development is only known for a limited number of species. Kutty et al. (2007) demonstrate that phytophagy in the form of leaf mining is the ancestral larval feeding habit for Scathophagidae and from phytophagy, two shifts to saprophagy and one shift to predation has occurred which a second origin of predation is from a saprophagous ancestor.

Altogether 157 species or subspecies are known to occur in Europe (Šifner 2008a, 2009a, b); the validity of further species needs to be revised in the future. In the present checklist, 77 species are listed from the Czech Republic (75 in Bohemia and 60 in Moravia) and 60 from Slovakia. The number of species known from individual European countries varies greatly and is dependent upon the level of faunistic and taxonomic research.

All the species from the Czech Republic and Slovakia can be reliably distinguishable by the structure of the male and female terminalia, and can be identified using the keys and figures by Šifner (2003a). Šifner (2003a) has compiled and summarised all the basic faunistic data. From the systematic point of view, two subfamilies, Scathophaginae and Delininae (Collin 1958), and eight tribes (Šifner 2003a) are recognised.


[1] Barták M. 2004: Faunistic records from the Czech and Slovak Republics. Diptera, Scathophagidae. In Bitušík P. (ed.): Dipterologica bohemoslovaca, Vol.12. Acta Facultatis Ecologicae 12 (Suppl. 1): 160.

[2] Barták M. & Kubík Š. 2006: Scathophagidae.  In Barták M. & Kubík Š. (eds): Diptera of Podyjí National Park and its Environs. Česká zemědělská Universita v Praze, pp. 362-365.

[3] Barták M. & Roháček J. 2009: Scathophagidae. In Roháček J. & Ševčík J. (eds): Diptera of the Poľana Protected Landscape Area – Biosphere Reserve (Central Slovakia). SNC SR, Administration of the PLA – BR Poľana, Zvolen, pp. 285-288.

[4] Bernasconi M.V., Valsangiacomo C., Piffaretti J.-C. & Ward P.I. 2000: Phylogeny of the Scathophagidae (Diptera, Calyptratae) Based on Mitochondrial DNA Sequences. Molecular Phylogenetics and Evolution 16: 308-315.

[5] Collin J.E. 1958: A short synopsis of the British Scatophagidae (Diptera). Transaction of the Society for British Entomology 13: 37-56.

[6] Kutty S. N., Bernasconi M.V., Šifner F. & Meier R 2007: Sensitivity analysis, molecular systematics and natural history evolution of Scathophagidae (Diptera: Cyclorrhaoha: Calyptratae). Cladistics 23: 64-83.

[7] Roháček J. & Barták M. 2006: Faunistic records. Scathophagidae. In Kinkorová J. (ed.): Dipterologica bohemoslovaca, Vol. 13. Acta Universitatis Carolinae, Biologica 50: 157.

[8] Šifner F. 2000: Three new species of the genus Scathophaga (Diptera, Scathophagidae). Folia Heyrovskyana 8: 193-196.

[9] Šifner F. 2003a: The family Scathophagidae (Diptera) of the Czech and Slovak Republics (with notes on selected Palaearctic taxa). Acta Musei Nationalis Pragae. Series B, Natural History 59(1-2): 1-90.

[10] Šifner F. 2003b: Two new species of the family Scathophagidae (Diptera) from the Czech and Slovak Republics. Journal of the National Museum, Natural History Series (Prague) 72: 77-80.

[11] Šifner F. 2004: Three new species of scathophagid flies from the Palaearctic region with a description of male Hexamitocera martineki (Diptera, Scathophagidae). Journal of the National Museum, Natural History Series (Prague) 173: 105-110.

[12] Šifner F. 2006a: Norellisoma jelineki sp. nov. – a new species from the Czech Republic (Diptera: Scathophagidae). Acta Entomologica Musei Nationalis Prague 46: 193-196.

[13] Šifner F. 2006b: Faunistic records from the Czech and Slovak Republics. Diptera, Scathophagidae. Entomofauna carpathica 18: 33.

[14] Šifner F. 2008a: A catalogue of the Scathophagidae (Diptera) of the Palaearctic region, with notes on their taxonomy and faunistics. Acta Entomologica Musei Nationalis Pragae 48(1): 111-196.

[15] Šifner F. 2008b: Three new species of the family Scathophagidae (Diptera) from the Palaearctic region with a redescription of the male of Amaurosoma longicorne. Acta Entomologica Musei Nationalis Pragae 48: 103-109.

[16] Šifner F. 2009a: Two new genera of Scathophagidae (Diptera). Acta Entomologica Musei Nationalis Pragae 49: 287-292.

[17] Šifner F. 2009b: Additions and corrections to the Catalogue of the Scathophagidae (Diptera) of  the Palaearctic Region. Acta Entomologica Musei Nationalis Pragae 49: 293-295.

Amaurosoma Becker, 1894
armillatum (Zetterstedt, 1846) CZ (B M ) SK
articulatum Becker, 1894 CZ (B M ) SK
bernasconii Šifner, 2008 CZ ( M )
brevifrons (Zetterstedt, 1838) CZ (B M ) SK
fasciatum (Meigen, 1826) CZ (B M )
flavipes (Fallén, 1819) CZ (B M ) SK
inerme Becker, 1894 CZ (B M ) SK
longicorne (von Roser, 1840) CZ (B )
leucostoma (Zetterstedt, 1846) CZ (B M )
minutum Becker, 1894 CZ (B M )
puberulum Becker, 1894 CZ (B M ) SK
tibiellum (Zetterstedt, 1838) CZ (B M ) SK
Gonatherus Rondani, 1856
planiceps (Fallén, 1819) CZ (B M ) SK
Orthacheta Becker, 1894
pilosa (Zetterstedt, 1838) CZ (B ) in ECV1 erroneously recorded for SK
Acerocnema Becker, 1894
macrocera (Meigen, 1826) CZ (B M ) SK
Cleigastra Macquart, 1835
apicalis (Meigen, 1826) CZ (B M ) SK
Gonarcticus Becker, 1894
abdominalis (Zetterstedt, 1846) CZ (B )
Hexamitocera Becker, 1894
martineki Šifner, 2003 CZ (B M ) SK
Jezekia Šifner, 2009
kmenti Šifner, 2009 CZ (B )
Megaphthalma Becker, 1894
pallida (Fallén, 1819) CZ (B M ) SK
Spathephilus Becker, 1894
breviventris (Loew, 1873) CZ (B M ) SK
Cordilura Fallén,1810
aemula (Collin, 1958) CZ (B M )
atrata (Zetterstedt, 1846) CZ (B M ) SK
ciliata (Meigen, 1826) CZ (B M ) SK
picipes (Meigen, 1826) CZ (B M ) SK
pubera (auct. nec Linnaeus, 1758) CZ (B M ) SK
pudica (Meigen, 1826) CZ (B M ) SK
rufimana (Meigen, 1826) CZ (B ) SK
umbrosa (Loew, 1873) CZ (B M ) SK
Norellisoma Wahlgren, 1917
alpestre (Schiner, 1864) CZ (B M ) SK
femorale (Loew, 1864) SK
flavicorne (Meigen, 1826) CZ (B M )
ivanae Šifner, 2003 SK
jelineki Šifner, 2006 CZ (B M )
lituratum (Meigen, 1826) CZ (B M ) SK
mirusae Šifner, 1974 CZ (B M ) SK
nervosum (Meigen, 1826) CZ (B M ) SK
spinimanum (Fallén, 1819) CZ (B M ) SK
striolatum (Meigen, 1826) CZ (B M ) SK
Parallelomma Becker, 1894
albipes (Fallén, 1819) CZ (B M ) SK
Phrosia Robineau - Desvoidy, 1830)
albilabris (Fabricius,1794) CZ (B M ) SK
Scoliaphleps Becker, 1894
ustulata (Zetterstedt, 1838) CZ (B M )
Gimnomera Rondani, 1867
castanipes (Becker, 1894) SK
cuneiventris (Zetterstedt, 1846) SK
slovaca Šifner, 2003 SK
tatrica Šifner, 2003 SK
Norellia Robineau - Desvoidy, 1830
spinipes (Meigen, 1826) CZ (B M ) SK
Cosmetopus Becker, 1894
dentimanus (Zetterstedt,1838) CZ (B )
Hydromyza Fallén, 1819
livens (Fabricius, 1874) CZ (B )
Chaetosa Cocquillett, 1898
punctipes (Meigen, 1826) CZ (B M ) SK
Paracosmetopus Hackman, 1956
helleni Hackman, 1956 CZ (B )
Pogonota Zetterstedt, 1860
barbata Zetterstedt, 1838 CZ (B )
Spaziphora Rondani, 1856
hydromyzina (Fallén, 1819) CZ (B M ) in ECV1 erroneously recorded for SK
Acanthocnema Becker, 1894
glaucescens (Loew, 1864) CZ (B M ) SK
nigrimana (Zetterstedt, 1846) CZ (B ) SK
Megaphthalmoides Gingdahl, 1936
unilineatus (Zetterstedt, 1838) CZ (B M ) SK
Microprosopa Becker, 1894
bartaki Šifner, 1999 CZ (B ) in ECV1 erroneously omitted [9]
haemorrhoidalis (Meigen, 1826) CZ (B M )
hoberlandti Šifner, 1981 CZ ( M ) SK
ozerovi Šifner, 2008 SK
pallidicauda (Zetterstedt, 1838) CZ (B M ) SK
Trichopalpus Rondani, 1856
fraternus Meigen, 1826) CZ (B M ) SK
Coniosternum Becker, 1894
lapponicum Ringdahl, 1920 CZ (B )
nelsoni Šifner, 2003 CZ (B )
obscurum (Fallén, 1819) CZ (B M ) SK
tinctinerve Becker, 1894 CZ (B )
Scathophaga Meigen, 1826  
bohemiae Šifner, 2000 CZ (B )
cineraria (Meigen, 1826) CZ (B M ) SK
furcata (Say, 1823) CZ (B M ) SK
inquinata (Meigen, 1826) CZ (B M ) SK
jizerensis Šifner, 2004 CZ (B )
lutaria (Fabricius, 1794) CZ (B M ) SK
pictipennis (Oldenberg, 1923) CZ (B ) SK
scybalaria (Linnaeus, 1758) CZ (B M ) SK in ECV1 erroneously omitted from SK [9]
suilla (Fabricius, 1794) CZ (B M ) SK
stercoraria (Linnaeus, 1758) CZ (B M ) SK
taeniopa (Rondani, 1867) CZ (B M )
vlastae Šifner, 2000 SK
Americina Malloch, 1923
media (Becker, 1894) CZ (B M ) SK
vittata (Meigen, 1826) CZ (B M ) SK
Delina Robineau-Desvoidy, 1830
anthrax (Schiner, 1864) CZ (B M ) SK
nigriceps (Becker, 1894) CZ (B M ) SK
nigrita (Fallén, 1819) CZ (B M ) SK
Leptopa Zetterstedt, 1838
filiformis Zetterstedt, 1838 CZ (B M ) SK in ECV1 erroneously omitted  [9]
Micropselapha Becker, 1894
filiformis (Zetterstedt, 1846) CZ (B M ) SK